A touch sensor substrate is equipped with: a base material having a first electrode placement surface, which is one example of a first surface; and first electrodes, which are one example of multiple electrodes positioned on the first surface. Each of the multiple first electrodes is equipped with a bottom surface in contact with the first surface, a top surface exposed from the first surface and facing the bottom surface, and side surfaces exposed from the first surface and connecting the bottom surface and the top surface, with the side surfaces and the bottom surface or top surface forming a blackening layer (BL).