The present disclosure relates to: an imaging element and an imaging-element drive method which enable negative-voltage fluctuation to be suppressed without increasing negative-voltage capacitance; an electronic device; and a program. A dual circuit is prepared for negative voltages. At times other than during exposure and reset, the negative voltage is supplied to readout transistors in lines other than a readout line. At timings other than these times, the negative voltage is supplied to the aforementioned readout transistors. Accordingly, even if the negative voltage fluctuates during exposure and reset, a stable negative voltage is supplied at timings at which other pixel data is read out, and thus fluctuation is suppressed. The present disclosure is applicable to imaging elements.