An image sensor may include an array of imaging pixels and row control circuitry. Each imaging pixel may include a photodiode, a floating diffusion region, a transfer transistor configured to transfer charge from the photodiode to the floating diffusion region, a dual conversion gain transistor coupled to the floating diffusion region, and a storage capacitor coupled to the dual conversion gain transistor. The capacitor may have a plate that receives a modulated control signal and the row control circuitry may be configured to modulate the control signal. To reduce image artifacts, the modulated control signal may be modulated low during the integration time of the pixel and may be modulated high during the high conversion gain readout time of the pixel.