Overrunning coupling and control assembly and control system for use therein are provided. The system includes a control member mounted for controlled shifting movement between the coupling faces of first of second coupling members. A one-way locking member such as a pawl is pivotally connected to the control member for movement between a disengaged position in which the control member is permitted to shift or rotate relative to the second coupling member and an engaged position between the control member and a locking member engaging portion of the second coupling member to lock the control member and the second coupling member together to prevent the control member from inadvertently shifting in a first direction relative to the second coupling member in the absence of an actuator command signal received by a bi-directional actuator subassembly including an output member connected to the control member.