A tool assembly for use with a closure actuator that is configured to move a first axial closure distance upon actuation thereof. A surgical end effector is operably coupled to a shaft assembly. A firing member assembly operably interfaces with an articulation member. A closure system operably interfaces with the closure actuator such that when the closure actuator is axially advanced through the first axial closure distance, the closure system disengages the firing member assembly and the articulation member and also moves a closure member axially through a second axial closure distance that is less than the first axial closure distance to apply closure motions to the end effector.