A radio station (2) transmits configuration information (501) to a radio terminal (1). The configuration information (501) indicates, on a cell-by-cell basis, at least one specific cell on which the radio terminal (1) is allowed to perform at least one of data transmission and data reception on a radio bearer used for uplink transmission or downlink transmission or both via a common Packet Data Convergence Protocol (PDCP) layer (402). As a result, in a radio architecture that provides tight interworking of two different Radio Access Technologies (RATs), it is for example possible to allow an eNB to indicate, to a radio terminal (UE), a specific cell on which the UE should perform uplink transmission.