A paint roller cover cleaner for cleaning a paint roller cover. The paint roller cover cleaner has one or more nozzles that spray the cover and that may each produce a fan-shaped jet in a plane that is at an angle to the longitudinal axis of the roller cover. The paint roller cover cleaner may also have a clamp for clamping a handle of the paint roller, another nozzle for cleaning an enclosure of the paint roller cover cleaner, a lid for the enclosure that drips within the mouth of the enclosure and does not move laterally across the roller cover to open. A system including a paint roller cover cleaner and a fluid supply that can contain the paint roller cover cleaner is also provided.