A multi-level signal receiver includes a data sampler circuit and a reference voltage generator circuit. The data sampler includes (M−1) sense amplifiers which compare a multi-level signal having one of M voltage levels different from each other with (M−1) reference voltages. The data sampler generates a target data signal including N bits, M is an integer greater than two and N is an integer greater than one. The reference voltage generator generates the (M−1) reference voltages, At least two sense amplifiers of the (M−1) sense amplifiers have different sensing characteristics.