An amplifier includes an amplification circuit, an equalization circuit, an output circuit, a first gain adjusting circuit, and a second gain adjusting circuit. The amplification circuit changes voltage levels of first and second amplification nodes based on first and second input signals. The equalization circuit changes the voltage levels of the first and second amplification nodes. The output circuit generates an output signal based on the voltage levels of the first and second amplification nodes. The first gain adjusting circuit changes voltage levels applied to the first and second amplification nodes based on the voltage levels of the first and second amplification nodes and a first gain control signal. The second gain adjusting circuit changes a voltage level of the output signal based on a second gain control signal.