An image sensor includes a pixel array unit in which pixels are arranged in a matrix, and multiple vertical signal lines provided in pixel column units. The multiple vertical signal lines provided in pixel column units are arranged in multiple wiring layers laminated on the pixel, and are arranged so that orthogonal projections of the vertical signal lines on the multiple wiring layers overlap. The wiring layer is provided with a connection portion for connecting the vertical signal line corresponding to the pixel to the pixel. A pixel signal is taken out from the vertical signal line through the connection portion.