A display substrate and a display device. A first opening region includes a first opening and a first opening peripheral region surrounding the first opening; a display region surrounds the first opening region the first signal lines extend along the first direction; the second signal lines extend along a second direction; the second signal lines pass through the first opening peripheral region along the second direction, each second signal line includes a longitudinal winding portion in the first opening peripheral region; the longitudinal winding portion partially surrounds the first opening; the longitudinal winding portion closest to the first opening among longitudinal winding portion of the plurality of second signal lines is an edge longitudinal winding portion, and the first floating electrode is in a same layer as the edge longitudinal winding portion and is at a side of the edge longitudinal winding portion close to the first opening.