A fuel cell stack that includes an actuating device or devices for selectively providing interdigitated reactant gas flow and straight reactant gas flow through reactant gas flow channels to reduce water accumulation in the diffusing media layers of the stack. In one embodiment, the fuel cell stack employs internal actuators that selectively close the inlet end of every other flow channel and the outlet end of every other opposite flow channel to provide the interdigitated flow. In another embodiment, the interdigitated flow is provided by external actuation where two inlet manifolds and two outlet manifolds are provided. One input manifold is closed to close the input ends of every other flow channel and one outlet manifold is closed to close the output ends of every other opposite flow channel.