A lens barrel which is capable of increasing the total length of the lens barrel in a shooting state even in a case where the collapsed lens barrel length is short, with a simple construction. A shooting optical system is disposed movably in an optical axis direction. A lens barrel driving member moves the shooting optical system in the optical axis direction while rotating around the optical axis. A lens barrel driving member moving unit moves the lens barrel driving member in the optical axis direction in a transition region during which the lens barrel moves between a shooting state and a non-shooting state while the driving member rotating.