An article of footwear that includes an outsole, including an outsole groove that extends generally in a thickness direction through the outsole to separate the outsole into first and second outsole elements. Furthermore, the article of footwear includes a midsole, including a midsole groove that extends generally in a thickness direction through the midsole to separate the midsole into first and second midsole elements. The first midsole element is operably secured to the first outsole element to define a first element assembly, and the second midsole element is operably secured to the second outsole element to define a second element assembly. Furthermore, the outsole groove and the midsole groove are substantially aligned with each other, and the first and second element assemblies interlock with each other.