An article of footwear is provided and includes a sole structure and an upper structure attached to the sole structure. The upper structure includes a vamp separated into medial and lateral portions, a tongue, and a lace. The tongue includes a main body portion and a first lace-retention feature attached to the main body portion. The main body portion has a first end attached to the medial and lateral portions, a second end that is moveable relative to the upper structure, and a longitudinal axis extending between the first end and the second end. The first lace-retention feature includes a first opening that receives the lace in a first direction substantially parallel to the longitudinal axis.