An article of footwear including heel portion movable relative to the forefoot portion from a first articulated configuration to a second articulated configuration is provided. An articulation assembly having a forefoot articulation member and a heel articulation member is also provided. The articulation assembly couples the forefoot portion to the heel portion and includes a hinge mechanism and a cam mechanism. The hinge mechanism may include a pin located within a socket. The cam mechanism may include a cam surface and a protrusion configured to ride on the cam surface. The article of footwear may include a locking mechanism having a first locking element that engages a second locking element in the first articulated configuration. The first locking element may be a first concavity formed in a surface and the second locking element may be a protrusion configured to extend into the first concavity in the first articulated configuration.