An applicator for applying a product to the eyelashes and/or the eyebrows including: a first application member including a first untwisted core, notably at least partially or even entirely made of synthetic material, and first application elements, a second application member including a second core twisted over at least a portion of the length of the first application member and second application elements supported by the second core, the first application member having, over at least a portion of its length, a spiralling induced by the twisting of the second core on the first application member.