An imaging device with (1) a substrate; (2) a substrate voltage supply that applies a first potential to the substrate during a light receiving period and applies a second potential to the substrate during a no-light receiving period; and (3) a plurality of pixels each including (a) a light conversion portion, (b) a storage portion that stores signal charges g from the light conversion portion when the first potential is applied to the substrate, (c) a first layer that is in the substrate and set apart from the storage portion by a predetermined distance and adjusts potential distribution in the substrate so that the signal charges generated in the light receiving portion when the second potential is applied to the substrate are swept to a rear surface side of the substrate, and (d) a vertical transfer portion that transfers a signal based on the signal charges in a vertical direction.