A liquid ejection device includes a first rolled medium accommodation unit, a second rolled medium accommodation unit and a liquid ejection head. The first rolled medium accommodation unit is configured to accommodate a rolled medium. The second rolled medium accommodation unit is configured to accommodate the rolled medium, the second rolled medium accommodation unit being movable. The liquid ejection head is configured to eject an ink as a liquid onto a medium supplied by unwound from the rolled medium accommodated in one of the first rolled medium accommodation unit and the second rolled medium accommodation unit.