A system and method for aligning and reducing the relative movement between adjacent fuel cells within a fuel cell stack. The inter-cell cooperation between fuel cells along a stacking dimension is enhanced by one or more datum placed along the edge of a bipolar plate that makes up a part of a cell-containing assembly. The datum is shaped along a thickness dimension that substantially coincides with the cell stacking dimension to promote a nested fit with a comparable datum on an adjacently-stacked bipolar plate. This nesting facilitates an interference fit that enhances the resistance to sliding movement between respective cells that may otherwise arise out of the occurrence of a significant acceleration along the dimension that defines the major surfaces of the plates, cells and their respective assemblies. In one form, the use of welding, bonding or related attachment of the datum to the plate promotes enhanced metallic support without the need for increasing the plate footprint and without having to overmold the datum directly onto the plate.