A filter element having a filter media support structure as well as a filter assembly having interacting filter media support structure components are disclosed. The support structure includes a support grid arranged on a clean side of the filter media. At least one support member is secured onto the support grid and extends outwardly away from the support grid and the filter media. The at least one support member is configured and adapted to supportively engage a respective mating support member to transfer forces from the filter element to the housing.